MD-PhD Class of 2016 - Harvard Medical School
Academic Degrees/PhD Titles/Postgraduate Year Destinations

Isha Agarwal
B.A., M.A., Harvard College (2009)
Ph.D., HSPH in Epidemiology� (2014)
DISSERTATION: Exploring the associations between diabetes, fibrosis-related biomarkers, and cardiovascular disease
M.D., Harvard Medical School (2016)

Tahir Ahmed
B.S., New York University (2006)
Ph.D., University of Cambridge (UK) in NIH-GPP (2013)
DISSERTATION: Selective Neuronal Loss, Microglial Activation and Adult Neurogenesis after Stroke: An in vitro and in vivo analysis of the SDF-1alpha/CXCR4 axis in focal ischemia and endogenous mechanisms of neural repair
M.D., Harvard Medical School (2016)
RESIDENCY: Anesthesiology at Beth Israel Deaconess Medical Center, Boston, MA

Milena M. Andzelm
B.A.., Harvard College (2006)
Ph.D., Harvard University in Immunology (2014)
DISSERTATION: Functional and genomic analysis of MEF2 transcription factors in neural development
M.D., Harvard Medical School (2016)
RESIDENCY: Child Neurology at Boston Children’s Hospital, Boston, MA

Alexander F. Bagley
B.S., B.S., Massachusetts Institute of Technology (2008)
Ph.D., Harvard University in Biophysics (2014)
DISSERTATION: Optically-Active Nanomaterials for Diagnostic and Therapeutic Applications in Ovarian Cancer
M.D., Harvard Medical School (2016)
RESIDENCIES: Preliminary Medicine at MD Anderson Cancer Center/University of Texas Medical School, Houston, TX
Radiation Oncology at MD Anderson Cancer Center, Houston, TX

Alexander G. Bick
B.S., M.S., Harvard College (2010)
Ph.D., Harvard University in Genetics(2014)
DISSERTATION: At the Heart of the Genome: Rare Genetic Variation, Cardiovascular Disease, and Therapy
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine at Massachusetts General Hospital, Boston, MA

Yamicia D. Connor
B.S., B.S., Massachusetts Institute of Technology (2007)
Ph.D., Massachusetts Institute of Technology in HST MEMP (2013)
DISSERTATION: Novel mechanisms of endothelial-epithelial interactions underlying cancer metastasis
M.D., Harvard Medical School (2016)
RESIDENCY: Ob/Gyn at Beth Israel Deaconess Medical Center, Boston, MA

Neir Eshel
B.A., Princeton University (2007)
MSc., University of College London in Clinical Neuroscience (2008)
Ph.D., Harvard University in Neuroscience (2014)
DISSERTATION: Arithmetic and local circuitry underlying dopamine prediction errors
M.D., Harvard Medical School (2016)
RESIDENCY: Psychiatry/Research Track at Stanford University Programs, Stanford, CA

Kay D. Everett
B.S., Massachusetts Institute of Technology (2007)
Ph.D., Harvard University in SEAS (2014)
DISSERTATION: Mechanisms and Implications of Fracture in Cardiovascular Stents
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital, Boston, MA

Jonathan D. Herman
Sc.B., Brown University (2007)
Ph.D., Harvard University in BBS (2014)
DISSERTATION: Halofuginone: A Story of How Target Identification and Multi-Step Evolution Informs Malaria Drug Discovery
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine/Research at New York-Presbyterian Hospital-Weill Cornell, New York, NY

Sarah J. Hill
A.B., Harvard College (2005)
MSc., Oxford University in Biochemistry (2006)
Ph.D., Harvard University in BBS-Genetics (2014)
DISSERTATION: New insights into BRCA1 function and its role in cancer development
M.D., Harvard Medical School (2016)
RESIDENCY: Anatomic Pathology at Brigham & Women’s Hospital, Boston, MA

Wen F. Hu
B.S., Yale University (2008)
Ph.D., Harvard University in Neuroscience (2014)
DISSERTATION: Building a Bigger Brain: Centriole Control of Cerebral Cortical Development
M.D., Harvard Medical School (2016)
RESIDENCIES: Preliminary Medicine at Brigham & Women’s Hospital, Boston, MA
Ophthalmology at Massachusetts Eye & Ear Infirmary, Boston, MA

Alison R. Hwong
Sc.B., Brown University� (2005)
Ph.D., Harvard University in Health Policy (Medical Sociology) (2016)
DISSERTATION: Social Ties and Health: An Analysis of Patient-Doctor Trust and Network-Based Public Health Interventions Through Randomized Experiments and Simulations
M.D., Harvard Medical School (2016)
RESIDENCY: Psychiatry at University of California, San Francisco, CA

David A. Kim
B.S., University of Toronto (2007)
Ph.D., Harvard University in Health Policy (Medical Sociology) (2015)
DISSERTATION: Social Networks and Health: From Epidemiology to Intervention
M.D., Harvard Medical School (2016)
RESIDENCY: Emergency Medicine at Stanford University Programs, Stanford, CA

Wendy W. Liu
A.B., Princeton University (2008)
Ph.D., Harvard University in Neuroscience (2014)
DISSERTATION: Dissecting Olfactory and Thermosensory Circuits in Drosophila
M.D., Harvard Medical School (2016)
RESIDENCIES: Preliminary Medicine at Beth Israel Deaconess Med Center, Boston, MA
Ophthalmology at Massachusetts Eye & Ear Infirmary, Boston, MA

Erin M. Loeliger
B.S., University of Maryland, Baltimore County (2005)
Ph.D., Harvard University in BBS-Genetics (2014)
DISSERTATION: Structure-Function Analysis of the Conserved Histone Chaperone Spt6
M.D., Harvard Medical School (2016)
RESIDENCY: Anesthesiology at Brigham & Women’s Hospital, Boston, MA

Karolina Maciag
A.B., Harvard College (2004)
Ph.D., Harvard University in Immunology (2014)
DISSERTATION: Effector functions and regulation of IFNy-induced immunity to intracellular pathogens
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine at Johns Hopkins Hospital, Baltimore, MD

Ilka A. Netravali
B.S. Princeton University (2003)
Ph.D. Harvard University in Immunology (2014)
DISSERTATION: Elucidation of plasmacytoid dendritic cell development
M.D., Harvard Medical School (2016)

Rishi V. Puram
B.S., B.S., Massachusetts Institute of Technology (2008)
Ph.D., Harvard University in BBS (2014)
DISSERTATION: Defining and Targeting Transcriptional Pathways in Leukemia Stem Cells
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine at Massachusetts General Hospital, Boston, MA

Aswin Sekar
B.S., University of California, Los Angeles (2007)
Ph.D., Harvard University in BBS (2014)
DISSERTATION: A natural allelic series of complex structural variants and its influence on the risk of lupus and schizophrenia
M.D., Harvard Medical School (2016)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital, Boston, MA

Jing (Meghan) Shan
B.S., Columbia University (2007)
Ph.D., Massachusetts Institute of Technology in HST MEMP� (2014)
DISSERTATION: High-throughput Approaches to Sourcing of Human Hepatocytes for Cell-based Therapies
M.D., Harvard Medical School (2016)
RESIDENCIES: Preliminary Medicine at Mt Auburn Hospital, Cambridge, MA
Ophthalmology at University of Southern California, Los Angeles, CA

Alexander O. Subtelny
A.B., Harvard College (2007)
Ph.D., Massachusetts Institute of Technology in Biology (2014)
DISSERTATION: mRNA poly(A)-tail lengths and their regulatory significance
M.D., Harvard Medical School (2016)
RESIDENCY: Pathology at Massachusetts General Hospital, Boston, MA

Veena Venkatachalam
B.S., Massachusetts Institute of Technology (2009)
Ph.D., Harvard University in Biophysics (2014)
DISSERTATION: Engineering microbial rhodopsins to expand the optogenetic toolkit
M.D., Harvard Medical School (2016)
RESIDENCIES: Preliminary Medicine at Brigham & Women’s Hospital, Boston, MA
Radiation Oncology/BWH-Harvard at Massachusetts General Hospital, Boston, MA