Dept. of Neurobiology
Harvard Medical School

220 Longwood Ave.
Boston, MA 02115-5701
office: (617) 432-1307
lab: (617) 432-3962
fax: (617) 432-1639





RT Born on PubMed or Google Scholar

*equal contributions


Born RT and Bencomo GM (2021) Illusions, Delusions, and Your Backwards Bayesian Brain: A Biased Visual Perspective. Brain Behav Evol. Mar 30:1-14. doi: 10.1159/000514859. Epub ahead of print. [link] PMID: 33784667.


Born RT (2019) Banishing “black/white thinking”: a trio of teaching tricks, eNeuro, 6(6):ENEURO.0456-19.2019 [link]. PMCID: PMC6900463

Born RT (2019) Unfriendly Skies for Medical Innovation, Wall Street Journal. op-ed, 17 November 2019 [pdf]

Born RT and Steinbusch HWM (2019) New methods for studying brain connectivity using viral tracing, J. Chem. Neuroanat., 102:101685. [pdf]

Hartmann TS, Raja S, Lomber SG and Born RT (2019) Cortico-cortical feedback from V2 exerts a powerful influence over the visually evoked local field potential and associated spike timing in V1. bioRxiv. doi:


*Lange RD, *Gómez-Laberge C, Haefner RM and Born RT (2018) Neural signatures of variable beliefs increase with task learning in V1, AREADNE, 20-24 June 2018, p. 76. [pdf]

György B, Meijer EJ, Ivanchenko MV, Tenneson K, Emond F, Hanlon KS, Indzhykulian AA, Volak A, Karavitaki KD, Tamvakologos PI, Vezina M, Berezovskii VK, Born RT, O'Brien M, Lafond JF, Arsenijevic Y, Kenna MA, Maguire CA, Corey DP (2018) Gene Transfer with AAV9-PHP.B Rescues Hearing in a Mouse Model of Usher Syndrome 3A and Transduces Hair Cells in a Non-human Primate. Mol Ther Methods Clin Dev. 13:1-13. [pdf] PMCID: PMC6297893


*Gómez-Laberge C, *Smolyanskaya A, Nassi JJ, Kreiman G and Born RT (2016) Bottom-up and top-down input augment the variability of cortical neurons, Neuron, 91:540-7. [pdf] PMCID: PMC5149417


Smolyanskaya A, Haefner RM, Lomber SG and Born RT (2015) A modality-specific feedforward component of choice-related activity in MT, Neuron, 87:208-19. [pdf] PMCID: PMC4512164

Trott AR and Born RT (2015) Input-gain control produces feature-specific surround suppression, J. Neurosci., 35(12):4973-82 [pdf]. PMCID: PMC4389596

Ruff DA and Born RT (2015) Gain modulation by feature attention in primate area MT depends on tuning strength, J. Neurophysiol., 113:1545-1555. [pdf] PMCID: PMC4346716

Mundell N, Beier K, Pan Y., Lapan S, Göz Aytürk D, Berezovskii VK, Wark A, Drokhlyansky E, Bielecki J, Born RT, Schier A, Cepko C (2015) Vesicular stomatitis virus enables gene transfer and transsynaptic tracing in a wide range of organisms, J. Comp. Neurol., 523(11):1639-63. [pdf] PMCID: PMC4458151

Stefan MI, Gutlerner JL, Born RT and Springer M (2015) The Quantitative Methods "Boot Camp": Teaching quantitative thinking and computing skills to graduate students in the life sciences, PLoS Comp. Biol., 11(4):e1004208 [pdf]. PMCID: PMC4399943

Nassi JJ, Cepko CC, Born RT and Beier K (2015) Neuroanatomy goes viral! Front. Neuroanat., 9:80. doi: 10.3389/fnana.2015.00080. PMCID: PMC4486834. [Abstract] [pdf]


Born RT, Trott AR and Hartmann TS (2014) Cortical magnification plus cortical plasticity equals vision? Vision Res., 111(Pt. B):161-9. [pdf] PMCID: PMC4400204

*Nassi JJ, *Gómez-Laberge C, Kreiman G and Born RT (2014) Cortico-cortical feedback increases the spatial extent of normalization, Front. Systems Neurosci., vol. 8, article 105, pp. 1-13. [pdf]. PMCID: PMC4039070


*Smolyanskaya A, *Ruff DA and Born RT (2013) Joint tuning for direction of motion and binocular disparity in macaque MT is largely separable, J. Neurophysiol., 110:2806-2816. [pdf]. PMCID: PMC3882821

Nassi JJ, Lomber SG and Born RT (2013) Cortico-cortical feedback contributes to surround suppression in V1 of the alert primate, J. Neurosci., 33(19):8504-8517. [pdf] PMCID: PMC3690087

Price NSC and Born RT (2013) Adaptation to speed in macaque MT/MST, J. Neurosci., 33(10):4359-4368. [pdf] PMCID: PMC3620717


Smolyanskaya A and Born RT (2012) Attention is more than meets the eye, Nature, 489(7416):371-2. [pdf].


Berezovskii VK, Nassi JJ and Born RT (2011) Segregation of feedforward and feedback projections in mouse visual cortex, J. Comp. Neurol., 519(18):3672-83. [pdf]. PMCID: PMC3219532

Ponce CR, Hunter JN, Pack CC, Lomber SG and Born RT (2011) Contributions of Indirect Pathways to Visual Response Properties in Macaque Area MT, J. Neurosci., 31(10):3894-903. [pdf] PMCID: PMC3117290

Hunter JN and Born RT (2011) Stimulus dependent modulation of suppressive influences in MT, J. Neurosci., 31(2):678-686. [pdf] PMCID: PMC3292058


Price NSC and Born RT (2010) Timescales of sensory- and decision-related activity in MT and MST, J. Neurosci., 30(42):14036-45. [pdf] PMCID: PMC3059109

Tsui JM, Hunter JN, Born RT and Pack CC (2010) The role of V1 surround suppression in MT motion integration, J. Neurophysiol., 103(6):3123-38. [pdf] PMCID: PMC2888240


Born RT, Tsui JM and Pack CC (2009) Temporal dynamics of motion integration, in Dynamics of Visual Motion Processing. Ilg, U. and Masson, G. eds. (New York: Springer). [pdf]

Price NSC and Born RT (2009) Representation of movement. Encyclopedia of Neuroscience Vol. 8, pp. 107-114 (Oxford: Elsevier Ltd.). [pdf]


Ponce CR and Born RT (2008) Stereopsis. Curr. Biol.. 18(18):R845-50. [pdf]

Ponce CR, Lomber SG and Born RT (2008) Integrating motion and depth via parallel pathways. Nat. Neurosci. 11(2):216-23. Epub 2008 Jan 13. [pdf] PMCID: PMC2377395

Pack CC and Born RT (2008) Cortical mechanisms for the integration of visual motion. in The Senses: A Comprehensive Reference. (Oxford: Elsevier Ltd. [pdf])


Roe AW, Parker AJ, Born RT, DeAngelis GC (2007) Disparity channels in early vision J Neurosci. 27(44):11820-11831 [pdf] PMCID: PMC2376798

Dauguet, J, Peled S, Berezovskii V, Delzescaux T, Warfield SK, Born R and Westin CF (2007) Comparison of fiber tracts derived from in-vivo DTI tractography with 3D histological neural tract tracer reconstruction on a macaque brain. NeuroImage 37(2):530-538. [pdf]


Pack CC, Conway BR, Born RT and Livingstone MS (2006) Spatiotemporal structure of nonlinear subunits in macaque visual cortex. J. Neurosci., 26:893-907. [pdf] PMCID: PMC1413500

Born RT, Pack CC, Ponce CR and Yi, Si. (2006) Temporal evolution of 2-dimensional direction signals used to guide eye movements. J. Neurophsyiol., 95:284-300. DOI, 10.1152/jn.01329.2004 [pdf]


Born RT and Bradley DC (2005) Structure and function of visual area MT. Annu. Rev. Neurosci., 28:157-89. DOI, 10.1146/annurev.neuro.26.041002.131052 [pdf]

Pack CC, Hunter JN and Born RT (2005) Contrast dependence of suppressive influences in cortical area MT of alert macaque. J. Neurophysiol., 93:1809-1815. [pdf]


Born RT (2004) Taking strategies to task. Neuron, 42:185-187. [pdf]

Pack CC, Gartland AJ and Born RT (2004) Integration of contour and terminator signals in visual area MT of alert macaque. J. Neurosci., 24:3268-3280. [pdf] PMCID: PMC6730032


Pack CC, Livingstone MS, Duffy KR and Born RT (2003) End-stopping and the aperture problem: Two-dimensional motion signals in macaque V1. Neuron, 39:671-680. [pdf]

(See also: van Wezel RJA and van der Smagt MJ (2003) Motion processing: How low can you go? Curr. Biol. 13:R840-R842. [pdf])

Pack CC, Born RT, and Livingstone MS (2003) Two-dimensional substructure of stereo and motion interactions in macaque visual cortex. Neuron, 37:525-535. [pdf]


Born RT and Pack CC (2002) Integration of motion cues for smooth pursuit eye movements. Ann. NY Acad. Sci., 956:453-455. [pdf]

Born RT, Pack CC, and Zhao R (2002) Integration of motion cues for the initiation of smooth pursuit eye movements. Prog. Brain Res., 140:225-237. [pdf]


Pack CC, Berezovskii VK and Born RT (2001) Dynamic properties of neurons in cortical area MT in alert and anesthetized macaque monkeys. Nature, 414:905-908. [pdf]

(For a different take on our results, presented by Movshon and colleagues, and our reply see: Cortical responses to visual motion in alert and anesthetized monkeys. Nature Neurosci., 6:3-4. [pdf])

Born RT (2001) Visual Processing: Parallel-er and Parallel-er. Curr. Biol. 11:R566-R568. [pdf]

Livingstone MS, Pack CC and Born RT (2001) 2-Dimensional substructure of MT receptive fields. Neuron 30:781-793. [pdf]

Pack CC and Born RT (2001) Temporal dynamics of a neural solution to the aperture problem in macaque visual area MT. Nature 409:1040-1042. [pdf]

(See also: Bradley, D (2001) MT Signals: Better with time. Nat. Neurosci. 4:346-348. [pdf])


Born RT (2000) Center-surround interactions in the middle temporal visual area of the owl monkey. J. Neurophysiol. 84(5):2658-2669. [pdf]

Born RT, Groh JM, Zhao R and Lukasewycz SL (2000) Segregation of object and background motion in visual area MT: Effects of microstimulation on eye movements. Neuron. 26(3):725-734. [pdf]

(See also: Treue, S and Ilg, U (2000) Image segmentation: A tug-of-war for the eyeball. Curr. Biol. 10:R746-R749. [pdf])

Berezovskii VK and Born RT (2000) Specificity of projections from wide-field and local motion processing regions within the middle temporal visual area of the owl monkey. J. Neurosci. 20(3):1157-1169. [pdf] PMCID: PMC6774190


Fung SH, Burstein D and Born RT (1998) In vivo microelectrode tip localization using magnetic resonance imaging. J. Neurosci. Methods, 80:215-224. [pdf] PMCID: PMC2972228


Geesaman BJ, Born RT, Andersen RA and Tootell RBH (1997) Maps of complex motion selectivity in the superior temporal cortex of the alert macaque monkey: A double-label 2-deoxyglucose study. Cerebral Cortex. 7:749-757. [pdf]

Groh JM, Born RT and Newsome WT (1997) How is a sensory map read out? Effect of microstimulation in area MT on saccades and smooth pursuit eye movements. J. Neurosci. 17(11):4312-4330. [pdf] PMCID: PMC6573560


Groh JM, Born RT and Newsome WT (1996) Interpreting sensory maps in visual cortex. IBRO News 24:11-12.


Tootell RBH, Reppas JB, Kwong KK, Malach R, Born RT, Brady TJ, Rosen BR and Belliveau JW (1995) Functional analysis of human MT and related cortical areas using magnetic resonance imaging. J. Neurosci. 15:3215-3230. [pdf] PMCID: PMC6577785


Born RT and Tootell RBH (1992) Segregation of global and local motion processing in primate middle temporal visual area. Nature 357:497-499. [pdf]


Born RT and Tootell RBH (1991) Spatial frequency tuning in single units of macaque striate cortex. Proc. Natl. Acad. Sci. USA 88:7066-7070. [pdf] PMCID: PMC52234

Born RT and Tootell RBH (1991) Single unit and 2-deoxyglucose studies of side inhibition in macaque striate cortex. Proc. Natl. Acad. Sci. USA 88:7071-7075. [pdf] PMCID: PMC52235


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