MD-PhD Class of 2013 - Harvard Medical School
Academic Degrees/PhD Titles/Postgraduate Year Destinations

Amma Fredua Agyemang
B.S., University of Albany (2002)
Ph.D., Harvard University in Immunology (2011)
ADVISOR: Dr. Michael Carroll
DISSERTATION: Regulation of Self-Reactive B cells: Implications forComplement and Toll-like Receptors��������������
M.D., Harvard Medical School (2013)
RESIDENCY: Postdoctoral Research at NIH, Bethesda, MD

Lauren Kaye Buhl (Barr)
B.A., M.B., University Pennsylvania (2004)
Ph.D., Massachusetts Institute of Technology in Brain and Cognitive�Sciences (2011)
ADVISOR: Dr. J. Troy Littleton
DISSERTATION: Synaptic Structure & Function at the DrosophilaLarval Neuromuscular Junction: A Molecular Analysis of Complexinand
Radish
M.D., Harvard Medical School (2013)
RESIDENCY: Preliminary Medicine at Lahey Clinic, Burlington, MA
Anesthesiology at Beth Israel Deaconess Medical Center, Boston, MA

Yiyin Erin Chen
B.A., University of Chicago (2006)
Ph.D., Massachusetts Institute of Technology in Biology (2011)
ADVISOR: Dr. Michael Laub
DISSERTATION: Phosphorylation-based control of cellular asymmetryand the cell cycle in Caulobacter crescentus
M.D., Harvard Medical School (2013)
RESIDENCY: Preliminary Medicine at Beth IsraelDeaconess Medical Center, Boston, MA and Dermatology at UCSF, San Francisco, CA

Yves Rene Chretien
B.A., Columbia University (2003)
Ph.D., Harvard University in Statistics (2010)
ADVISOR: Dr. Carl N. Morris and Dr. Donald B. Rubin
DISSERTATION: A New Kullback-Leibler Statistic to QuantifyDiscrimination and Calibration in Binomial Models
M.D., Harvard Medical School (2013)
RESIDENCY: Preliminary Medicine/Radiology atMassachusetts General Hospital, Boston, MA

Sonia Cohen
B.A., Johns Hopkins University (2002)
Ph.D., Harvard University in BBS-Genetics (2011)
ADVISOR: Dr. Michael Greenberg
DISSERTATION: Activity-Dependent Phosphorylation of MeCP2 inNeural Development and Rett Syndrome
M.D., Harvard Medical School (2013)
RESIDENCY: General Surgery at Massachusetts General Hospital, Boston, MA

Joseph Wang Franses
B.S., Purdue University (2005)
Ph.D., Massachusetts Institute of Technology in HST MEMP/Chemical Engineering (2011)
ADVISOR: Dr. Elazer Edelman
DISSERTATION: Regulatory Roles of Endothelial Cells in Cancer
M.D., Harvard Medical School (2013)
RESIDENCY: Internal Medicine at Massachusetts General Hospital,Boston, MA

Daniel Steven Herman
B.S., Massachusetts Institute of Technology (2004)
Ph.D., Harvard University in BBS-Genetics (2011)
ADVISOR: Dr. Jonathan Seidman
DISSERTATION: Massively parallel, targeted DNA resequencing andcardiovascular disease
M.D., Harvard Medical School (2013)
RESIDENCY: Pathology at University of WashingtonAffiliated Hospitals, Seattle, WA

Marie Alice Hollenhorst
B.S., Stanford University (2005)
Ph.D., Harvard University in Chemical Biology (2011)
ADVISOR: Dr. Christopher T. Walsh
DISSERTATION: Biosynthesis and biological activity of thedapdiamide antibiotics
M.D., Harvard Medical School (2013)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital,Boston, MA

Peggy Ping Hsu
A.B., Princeton University (2003)
Ph.D., Massachusetts Institute of Technology in Biology (2011)
ADVISOR: Dr. David M. Sabatini
DISSERTATION: The Identification of the mTOR-RegulatedPhosphoproteome and a Mediator of Feedback Inhibition to PI3K-Akt
M.D., Harvard Medical School (2013)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital,Boston, MA

Jeffrey Joseph Ishizuka
B.A., Williams College (2004)
D.Phil., NIH-GPP in U Oxford (UK) (2008)
ADVISOR: Dr. Andrew McMichael and Dr. Jonathan W. Yewdell
DISSERTATION: Immunodominance and Interconnection in theCD8+ T-cell Response to Influenza and other Viruses
M.D., Harvard Medical School (2013)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital,Boston, MA

Robert Mawunyo Koffie
B.S., B.S., Indiana University (2007)
Ph.D., Harvard University in Biophysics (2011)
ADVISOR: Dr. Bradley T. Hyman
DISSERTATION: Molecular Imaging of Synaptic Changes inAlzheimer’s Disease
M.D., Harvard Medical School (2013)
RESIDENCY: Neurological Surgery at Massachusetts General Hospital,Boston, MA

Thomas Michael Kolokotrones
A.B., Harvard College (2000)
M.P.H., Harvard School of Public Health (2013)
Ph.D., Harvard University in BBS-Cell Biology (2010)
ADVISOR: Dr. Walter Fantana
DISSERTATION: Curvature in Metabolic Scaling
M.D., Harvard Medical School (2013)
RESIDENCY: Transitional at Harbor-UCLA Medical Center,Los Angeles, CA

Peter Grant Miller
B.S., Massachusetts Institute of Technology (2005)
Ph.D., Harvard University in Immunology (2011)
ADVISOR: Dr. Benjamin Ebert
DISSERTATION: Targeting Leukemia Stem Cells in the HematopoieticNiche
M.D., Harvard Medical School (2013)
RESIDENCY: Internal Medicine at Brigham & Women’s Hospital,Boston, MA

Mobolaji Oluyemisi Olurinde
B.S., B.S., University of Florida (2002)
S.M., Massachusetts Institute of Technology in Biological Engineering (2007)
Ph.D., Massachusetts Institute of Technology in HST�MEMP/Biophysics (2010)
ADVISOR: Dr. Jianzhu Chen
DISSERTATION: Factors Contributing to T Cell Persistence in aTolerizing Tumor Environment
M.D., Harvard Medical School (2013)
RESIDENCY: Anesthesiology at Penn State, Hershey, PA

Vikram Pattanayak
B.A., M.S., University Pennsylvania (2004)
Ph.D., Harvard University in Chemistry (2011)
ADVISOR: Dr. David R. Liu
DISSERTATION: Revealing the DNA Cleavage Specificities of Therapeutically Relevant Endonucleases Through In Vitro Selection
M.D., Harvard Medical School (2013)
RESIDENCY: Pathology at Massachusetts General Hospital, Boston,MA

Sidharth Venkata Puram
B.S., B.S., Massachusetts Institute of Technology (2005)
Ph.D., Harvard University in BBS-Pathology (2011)
ADVISOR: Dr. Azad Bonni
DISSERTATION: Calcium-Regulated Ubiquitin Signaling at theCentrosome Drives Dendrite Patterning in the Mammalian Brain
M.D., Harvard Medical School (2013)
RESIDENCY:Preliminary Surgery at Massachusetts General Hospital, Boston, MA
Otolaryngology - Head and Neck Surgery at Massachusetts Eye and Ear Infirmary, Boston, MA

Benjamin Isaac Rapoport
A.B., A.M., Harvard College (2003)
M.Sc., Oxford University in Mathematics (2004)
Ph.D., Massachusetts Institute of Technology in Electrical Engineering and Computer Science (2011)
ADVISOR: Dr. Rahul Sarpeshkar
DISSERTATION: Glucose-Powered Neuroelectronics
M.D., Harvard Medical School (2013)
RESIDENCY: Neurological Surgery at New York-Presbyterian�Hospital-Weill�Cornell�Medical�Center, New York, NY

Narie Yoo Storer
A.B., Harvard College (2002)
Ph.D., Harvard University in BBS-Genetics (2011)
ADVISOR: Dr. Leonard I. Zon
DISSERTATION: Characterization of the myogenic program inzebrafish rhabdomyosarcoma
M.D., Harvard Medical School (2013)
RESIDENCY: Pediatrics at Children’s Hospital, Boston, MA

Christos George Tsokos
B.S., Cornell University (2004)
Ph.D., Massachusetts Institute of Technology in Biology (2011)
ADVISOR: Dr. Michael Laub
DISSERTATION: Regulation of Cell Fate Asymmetry in Caulobacter�crescentus by a Complex of Two Component Signaling Proteins
M.D., Harvard Medical School (2013)
RESIDENCY: Pathology at UCSF, San Francisco, CA

Rochelle Marie Witt
B.S., B.S., University of Illinois Urbana-Champaign (1995)
Ph.D., Harvard University in Neuroscience (2010)
ADVISOR: Dr. Rosalind A. Segal
DISSERTATION: Of Structure and Function: The Nature of the SonicHedgehog-Heparan Sulfate Proteoglycan Interaction
M.D., Harvard Medical School (2013)
RESIDENCY: Child Neurology at NYP Hosp-Weill Cornell MedicalCenter, New York, NY